The Complexe des lacs Ambondro et Sirave (Complex of Lakes Ambrondo and Sirave, or CLAS) features a variety of habitats including dune lakes, mangrove forests, intertidal zones and marshes. It supports a remarkable diversity of species such as the endangered Verreaux’s sifaka (Propithecus verreauxi) and ring-tailed lemur (Lemur catta), and the near-threatened brown lemur (Eulemur fulvus). It also supports birds including the endangered Madagascar heron Ardea humbloti, Madagascar pond-heron Ardeola idae and Madagascar teal Anas bernieri, as well as the vulnerable black-banded plover Charadrius thoracicus, and is an important breeding and nesting ground for migratory waterbirds such as the curlew sandpiper Calidris ferruginea, common greenshank Tringa nebularia and the common ringed plover Charadrius hiaticula. Other species depending on the Site include the greater flamingo, which contributes greatly to its tourism value. The Site hosts the largest mangrove forest in the region and nourishes about 90% of the local population; human activities include agriculture, fishing and hunting.